Davies has poached The Observer's ad manager Chris Willis to replace Rob Atkinson, Mail on Sunday's head of trading.
Atkinson has been promoted to a senior role at sister Associated Newspapers title Ireland on Sunday.
Atkinson's move to the Irish title follows Associated's decision to bring Davies back to the Mail on Sunday from his ad director post in Ireland. He replaced Sue Dear at the Mail on Sunday two weeks ago.
Davies was unavailable to comment.
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