Sales of The Independent continued to rise following its decision to launch a tabloid version. In February, its year-on-year sales, excluding bulk copies, were up 20 per cent to 222,799.
The Times is selling more copies of its tabloid version than The Independent, but the overall impact on its circulation has been smaller. Its sales were down 3 per cent to 614,610 in February.
Increased sales of The Independent seem to have had the greatest impact on sales of The Guardian, which are down 10.5 per cent to 352,005. The title said last month that it would not be publishing a tabloid version.
The Daily Telegraph is also suffering, with a drop of 4 per cent in February to 873,380, continuing its recent pattern of decline.
In the red-top market, the Daily Star continued to perform well, up 8 per cent to 909,240, while both The Sun and the Daily Mirror continued to lose sales. The Sun's circulation was down 3 per cent to 3,397,372 and the Daily Mirror's down 7 per cent to 1,900,250.
The mid-market titles also appear to have been squeezed by The Times' and The Independent's tabloid variants. Sales of the Daily Mail were down 3 per cent to 2,311,849, although the Daily Express' sales rose by 2,000 copies to 906,738.
Sales of quality Sunday titles were also hit, with The Sunday Times, The Sunday Telegraph and The Observer all losing readers.