Both quality and red-top titles were hit as the downward trend in circulations continued. The performance in the quality market was less dismal, with the FT's circulation rising by 3.4 per cent to 461,033 and The Daily Telegraph down by just 0.6 per cent to 896,197.
However, The Times' circulation was down by 4.5 per cent to 639,547 and The Guardian's by 3.6 per cent to 366,233. The Independent was down 1.2 per cent to 252,714.
Among the mid-market titles, the Daily Mail was down by 2.3 per cent to 2,342,915, while its rival the Daily Express performed even worse, down 8.8 per cent to 758,753.
Sales of the Daily Mirror fell by 5.6 per cent and The Sun was down by 3.7 per cent to 3,031,724.
Sales of all Sunday titles fell, with The Sunday Times down by 8.8 per cent to 1,236,911 and the News of the World by 4.3 per cent to 3,308,111.
While some free newspapers continued to record circulation increases, sales of the Evening Standard fell by 17.7 per cent to 266,575. News International's thelondonpaper saw its circulation rise slightly on the previous month to 502,158, while Associated Newspapers' London Lite fell by 0.14 per cent to 400,212.
Metro, Associated's morning freesheet, rose by 0.6 per cent to 1,125,651.